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Martín-Hervás, M. R., Carmona, L., Krug, P. J., Gosliner, T., Cervera, J. L. & Malaquias, M. A. E. 2025. Origin and Biogeography of the Colourful Sap-Sucking Sea Slugs Genus Thuridilla Bergh, 1872 (Mollusca, Gastropoda, Heterobranchia). Journal of Biogeography, e15171. 

Bharate, M., Turani, M., Uribe, J. E., & Malaquias, M. A. E. 2025. On the relationships and origin of the worldwide Haminoea sensu lato marine molluscs (Gastropoda, Cephalaspidea) and the Importance of in‐depth taxon sampling. Zoologica Scripta.

Malaquias, M. A. E., Turani, M., Lopes, E., & Cervera, J. L. 2025. The Heterobranchia gastropods from the Cabo Verde Islands (NE Atlantic): a hot-spot of endemism. Regional Studies in Marine Science, 86: 104193.

Ekimova, I. A., Nikitenko, E. D., Stanovova, M. V., Antokhina, T. I., Schepetov, D. M., & Malaquias, M. A. E. 2025. A deceptive variety: integrative systematics and phylogeography suggest Adalaria proxima and Adalaria loveni represent a single species (Gastropoda: Nudibranchia). Systematics and Biodiversity, 23(1), 2468639.

Barreiros, J. P., Botelho, A. Z., Costa, A. C., Parente, M. I., Porteiro, F. M., Barcelos, L. M. D., Azevedo, J., Barreiros, J. B., Menezes, G. M., Lhoumeau, S., Malaquias, M., Coelho, R., Prestes, A. C. L., Faria, J., Torres, P., Albuquerque, I., Ramos, J. B., Fontes, J., Correia, L. A. & Luz, R. 2024. Guia práctico da flora e fauna marinha dos Açores / Filed guide of Azorean marine flora and fauna. Instituto Açoreano de Cultura  (IAC), Angra do Heroismo, 528p. [book]

Malaquias, M. A. E. 2024. Nudibranchs and other sea slugs of the Azores. Marine gastropods. Letras Lavadas, Ponta Delgada, Açores, (ISBN: 9789897355387), 140p. [book]

Lobo-da-Cunha, A., Bartolomaeus, T. & Malaquias, M. A. E. 2024. Euopisthobranchs: Musculature. In: Starck, J. M. (Ed.), Microscopic Anatomy of Animals. John Wiley & Sons, p. 1–17. [Encyclopaedia chapter]

Lobo-da-Cunha, A., Bartolomaeus, T. & Malaquias, M. A. E. 2024. Euopisthobranchs: Integument. In: Starck, J. M. (Ed.), Microscopic Anatomy of Animals. John Wiley & Sons, p. 1–7. [Encyclopaedia chapter]

Grishina, D. Y., Schepetov, D. M., Antokhina, T. I., Malaquias, M. A. E., Valdés, Á., & Ekimova, I. A. 2024. Panmixia and local endemism: a revision of the Eubranchus rupium species complex with a description of new species. Invertebrate Systematics, 38: IS24032.

Ekimova, I. A., Schepetov, D. M., Green, B., Stanovova, M. V., Antokhina, T. I., Gosliner, T., Malaquias, M. A. E. & Valdés, Á. 2024. Scaling the high latitudes: evolution, diversification, and dispersal of Coryphella nudibranchs across the Northern Hemisphere. Molecular Phylogenetics and Evolution, 201: 108214.

Siegwald, J., & Malaquias, M. A. E. 2024. Bringing light into deep-sea biodiversity: a systematic revision and molecular phylogeny of the genus Scaphander Montfort, 1810 (Gastropoda: Cephalaspidea), with a focus on the Indo-Pacific. Zoological Journal of the Linnean Society, zlad201.

Lobo-da-Cunha, A., Bartolomaeus, T. & Malaquias, M. A. E. 2023. Euopisthobranchs: Intestinal System. In: Starck, J. M. (Ed.), Microscopic Anatomy of Animals. John Wiley & Sons, p. 1–24. [Encyclopaedia chapter]

Lobo-da-Cunha, A., Bartolomaeus, T. & Malaquias, M. A. E. 2023. Euopisthobranchs: Introduction. In: Starck, J. M. (Ed.), Microscopic Anatomy of Animals. John Wiley & Sons, p. 1–9. [Encyclopaedia chapter]

Lobo-da-Cunha, A., Bartolomaeus, T. & Malaquias, M. A. E. 2023. Euopisthobranchs: Respiratory System. In: Starck, J. M. (Ed.), Microscopic Anatomy of Animals. John Wiley & Sons, p. 1–8. [Encyclopaedia chapter]

Lobo-da-Cunha, A., Bartolomaeus, T. & Malaquias, M. A. E. 2023. Euopisthobranchs: Skeletal/Support System. In: Starck, J. M. (Ed.), Microscopic Anatomy of Animals. John Wiley & Sons, p. 1–7. [Encyclopaedia chapter]

Bartolomaeus, T., Malaquias, M. A. E. & Lobo-da-Cunha, A. 2023. Euopisthobranchs: Body Cavities. In: Starck, J. M. (Ed.), Microscopic Anatomy of Animals. John Wiley & Sons, p. 1–10. [Encyclopaedia chapter]

Bartolomaeus, T., Malaquias, M. A. E. & Lobo-da-Cunha, A. 2023. Euopisthobranchs: Circulatory System. In: Starck, J. M. (Ed.), Microscopic Anatomy of Animals. John Wiley & Sons, p. 1–9. [Encyclopaedia chapter]

Bartolomaeus, T., Malaquias, M. A. E. & Lobo-da-Cunha, A. 2023. Euopisthobranchs: Excretory System. In: Starck, J. M. (Ed.), Microscopic Anatomy of Animals. John Wiley & Sons, p. 1–11. [Encyclopaedia chapter]

Landau, B., Harzhauser, M., Malaquias, M. A. E. 2023. The Pliocene Gastropoda (Mollusca) of Estepona, southern Spain. Part 22: Marine Heterobranchia (excluding Pyramidelloidea). Cainozoic Research, 23(2): 337–432.

Turani, M., Valdés, Á., and Malaquias, M. A. E. 2023. Molecular phylogeny of the marine snail genus Haminoea (Gastropoda, Cephalaspidea): a framework to study marine diversity and speciation. Zoologica Scripta, 53(1): 52–77.

Tibiriçá, Y., Pola, M., Pittman, C., Gosliner, T. M., Malaquias, M. A., and Cervera, J. L. 2023. A Spanish dancer? No! A troupe of dancers: a review of the family Hexabranchidae Bergh, 1891 (Gastropoda, Heterobranchia, Nudibranchia) A troupe of Spanish dancer. Organisms Diversity & Evolution, 23: 697–742.

Ekimova, I. A., Nikitenko, E., Stanovova, M. V., Schepetov, D. M., Antokhina, T. I., Malaquias, M. A. E., and Valdés, Á. 2023. Unity in diversity: morphological and genetic variability, integrative systematics, and phylogeography of the widespread nudibranch mollusc Onchidoris muricata. Systematics and Biodiversity, 21(1), 2246472.

Malaquias, M. A. E. and Afonso, P. 2023. An account on the marine Heterobranchia gastropods collected during the Mission “GAMPA Graciosa 2014” (Archipelago of the Azores, mid-north Atlantic). Arquipelago. Life and Marine Sciences, 38: 43–49.

Campanyà-Llovet, N., Bates, A. Cuvelier, D., Giacomello, E., Catarino, D., Gooday, A. J., Berning, B., Figuerola, B., Malaquias, M. A. E., Moura, C. J., Xavier, J. R., Sutton, T. T., Fauconnet, L., Ramalho, S. P., Neves, B. d. M., Machado, G. M., Horton, T., Gebruk, A.V., Minin, K., Bried, J., Molodtsova, T., Silva, M. A., Dilman, A., Kremenetskaia, A., Costa, E. F. S., Clarke, J., Martins, H. R., Pham, C. K., Carreiro-Silva, M., and Colaço, A. 2023. FUN Azores: a FUNctional trait database for the meio-, macro-, and megafauna from the Azores Marine Park (Mid-Atlantic Ridge). Frontiers in Ecology and Evolution, 11: 1050268.

 Valdés, Á., Feliciano, K., and Malaquias, M. A. 2023. The genus Pupa Röding, 1798 (Mollusca, Gastropoda, Acteonidae) in New Caledonia with notes on Recent species. Zootaxa, 5270(3), 471–506.

Araújo, A. K., Pola, M., Malaquias, M. A. E., Vitale, F., and Cervera, J. L. 2023. Integrative taxonomy reveals that not all European reddish runcinids are the same: the case of the Runcina ferruginea Kress, 1977 (Gastropoda, Heterobranchia, Runcinida) species-complex, with the description of a new genus. Invertebrate Systematics, 37(1), 61–77.

Fassio, G., Stefani, M., Russini, V., Buge, B., Bouchet, P., Treneman, N., Malaquias, M. A. E., Schiaparelli, S., Modica, M. V., Oliverio, M. 2023. Neither slugs nor snails: a molecular reappraisal of the gastropod family Velutinidae, Zoological Journal of the Linnean Society, 197(4), 924–964.

Oskars, T. R., and Malaquias, M. A. E. 2022. Systematic revision of the Indo-West Pacific bubble-snails of the genus Haloa (Pilsbry, 1921) (Cephalaspidea: Haminoeidae). Invertebrate Systematics, 36(5): 436–492.

Ekimova, I., Valdés, Á., Malaquias, M. A. E., Rauch, C., Chichvarkhin, A., Mikhlina, A., Antokhina, T., Chichvarkhina, O., and Schepetov, D. 2022. High-level taxonomic splitting in allopatric taxa causes confusion downstream: a revision of the nudibranch family Сoryphellidae. Zoological Journal of the Linnean Society, 196, 215–249.

Hellem, E., and Malaquias, M. A. E. 2022. The taxonomic status of the headshield slug genus Nakamigawaia Kuroda and Habe, 1961 (Gastropoda: Cephalaspidea: Aglajidae), with the description of a new species from the Western Pacific. Journal of Natural History, 55(35–36): 2231–2244.

Siegwald, J., Kano, Y., and Malaquias, M. A. E. 2022. A global phylogeny of the deep-sea gastropod family Scaphandridae (Heterobranchia: Cepahalaspidea): redefinition and generic classification. Molecular Phylogenetics and Evolution, 169: 107415.

Siegwald, J. and Malaquias, M. A. E. 2022. Revisiting the deep-sea Atlantic gastropod species Scaphander gracilis Watson, 1883: First data on its morphology, systematics, and ecology. Journal of Natural History, 55(47–48): 3053–3066.

Ramirez-Llodra, E., Pedersen, T., Dexter, K.F., Hauquier, F., Guilini, K., Mikkelsen, N., Borgersen, G., Van Gyseghem, M., Vanreusel, A. and Vilas, D., 2021. Community structure of deep fjord and shelf benthic fauna receiving different detrital kelp inputs in northern Norway. Deep Sea Research Part I: Oceanographic Research Papers, 168, p.103433. DOI: .

Neuhaus, J., Rauch, C., Bakken, T., Picton, B., Pola, M. and Malaquias, M. A. E. 2021. The genus Jorunna (Nudibranchia: Discodorididae) in Europe: a new species and a possible case of incipient speciation. Journal of Molluscan Studies, 87(4), eyab028. DOI: . 

Feliciano, K., Malaquias, M. A. E., Stout, C., Brenzinger, B., Gosliner, T. M., and Valdés, Á. 2021. Molecular and morphological analyses reveal pseudocryptic diversity in Micromelo undatus  (Bruguière, 1792) (Gastropoda: Heterobranchia: Aplustridae). Systematics and Biodiversity, XX: 1-25. . 

Araújo, A. K., Pola, M., Malaquias, M. A. E., Ballesteros, M., and Cervera, J. L. 2021. Molecular phylogeny of European Runcinida (Gastropoda, Heterobranchia): the discovery of an unexpected pool of species complex, with special reference to the case of Runcina coronata (Quatrefages, 1844). Zoological Journal of the Linnean Society, XX: 1-28. 

Martín-Hervás, M. R., Carmona, L., Malaquias, M. A. E., Krug, P. J., Gosliner, T. M. and Cervera, J. L. 2021. A molecular phylogeny of Thuridilla Bergh, 1872 sea slugs (Gastropoda, Sacoglossa) reveals a case of flamboyant and cryptic radiation in the marine realm. Cladistics, 37: 647–676.

Varney, R. M., Brenzinger, B., Malaquias, M. A. E., Meyer, C. P., Schrödl, M. and Kocot, K. 2021.  Assessment of mitochondrial genomes for heterobranch gastropod phylogenetics. BMC Ecology and Evolution, 21(1): 1-14. 

Bharate, M., Apte, D., and Malaquias, M. A. E. 2020. A massive aggregation of the mangrove snail Bakawan rotundata (A. Adams, 1850) (Cephalaspidea: Haminoeidae) in India. Regional Studies in Marine Science, 39: 101406. . 

Sørensen, C. G., Rauch, C., Pola, M. and Malaquias, M. A. E. 2020. Integrative taxonomy reveals a cryptic species of the nudibranch genus Polycera (Polyceridae) in European waters. Journal of the Marine Biological Association of the United Kingdom, 100(5): 733-752. .

Oskars, T. R. & Malaquias, M. A. E. 2020. The temperate Australasian genus Papawera Oskars and Malaquias, 2019 (Gastropoda: Cephalaspidea: Haminoeidae), with a redescription of P. zelandiae and P. maugeansis. Journal of Natural History, 54(21-22): 1343-1362. .

Oskars, T. R. & Malaquias, M. A. E. 2020. Systematic revision of the Indo-West Pacific colourful bubble-snails of the genus Lamprohaminoea Habe, 1952 (Cephalaspidea : Haminoeidae). Invertebrate Systematics, 34: 727–756. .

Oskars, T. R. & Malaquias, M. A. E. 2020. Systematic revision of the Indo-West Pacific mangrove-associated snails of the genus Bakawan (Cephalaspidea: Haminoeidae). Journal of Molluscan Studies, 86: 323–341. 

Lobo-da-Cunha, A., Alves, Â., Oliveira, E., Wranik, W., and Malaquias, M. A. E. 2020. The copulatory apparatus of the marine gastropod Haminella solitaria (Say, 1822) (Heterobranchia: Cephalaspidea) and its phylogenetic relevance. Invertebrate Biology, 139(1): e12281. 

Siegwald, J., Pastorino, G., Oskars, T., and Malaquias, M. A. E. 2020. A new species of the deep-sea genus Scaphander (Gastropoda, Cephalaspidea) from the Mar del Plata submarine canyon off Argentina. Bulletin of Marine Sciences, 96(1):111–126.&�Բ�����;

Kocot, K. M., Todt, C., Mikkelsen, N. T., & Halanych, K. M. 2019. Phylogenomics of Aplacophora (Mollusca, Aculifera) and a solenogaster without a foot. Proceedings of the Royal Society B, 286(1902), 20190115. DOI: .

Oskars, T. R. and Malaquias, M. A. E. 2019. A molecular phylogeny of the Indo-West Pacific species of Haloa sensu lato gastropods (Cephalaspidea: Haminoeidae): Tethyan vicariance, generic diversity, and ecological specialization. Molecular Phylogenetics and Evolution, 139: 

Araujo, A. K., Pola, M., Malaquias, M. A. E., Cervera, J. L. 2019. To be or not to be? What molecules say about Runcina brenkoae Thompson, 1980 1 (Gastropoda: Heterobranchia: Runcinida). Sciencia Marina, 83(3): 223-235. .

Oskars, T. R., Too, C. C., Rees, D., Mikkelsen, P., Willassen, E., Malaquias, M. A. E. 2019. A molecular phylogeny of the gastropod family Haminoeidae sensu lato (Heterobranchia, Cephalaspidea), - a generic revision. Invertebrate Systematics, 33(2): 426-472. .

Aslam, S., Oskars, T. R., Siddiqui, G., Malaquias, M. A. E. 2019. Beyond shells: First detailed morphological description of the mangrove-associated gastropod Haminoea aff.fusca (A. Adams, 1850) (Cephalaspidea: Haminoeidae), with a COI phylogenetic analysis. Zoosystema, 41(16): 313-326. .

Mikkelsen, N. T., Todt, C., Kocot, K. M., Halanych, K. M., Willassen, E. 2019. Molecular phylogeny of Caudofoveata (Mollusca) challenges traditional views.Molecular Phylogenetics and Evolution 132:&�Բ�����;138–150.&�Բ�����;

Moles, J., Avila, C., Malaquias, M. A. E. 2019. Unmasking Antarctic mollusc lineages: novel evidence from philinoid snails (Gastropoda: Cephalaspidea). Cladistics, 35:1–27. 

Mikkelsen, N. T., Kocot, K. M., Halanych, K. M. 2018. Mitogenomics reveals phylogenetic relationships of caudofoveate aplacophoran molluscs. Molecular Phylogenetics and Evolution127. 429–436. 

Mikkelsen, N. T., Todt, C. 2018. One or many? Molecular versus morphological diversity in the aplacophoran Chaetoderma nitidulumLovén, 1844 (Mollusca: Caudofoveata). Journal of Molluscan Studies84. 113–131. 

Bharate, M., Oskars, T. R., Narayana, S., Ravinesh, R., Kumar, A. B., Malaquias, M. A. E. 2018. Description of a new species of Haminoea (Gastropoda: Cephalaspidea) from India, with an account of the diversity of the genus in the Indo-West Pacific. Journal of Natural History, 52 (37–38): 2437–2456. 

Austin, J., Gosliner, T., Malaquias, M. A. E. 2018. Systematic revision, diversity patterns, and trophic ecology of the tropical Indo-West Pacific sea slug genus PhanerophthalmusA. Adams, 1850 (Cephalaspidea, Haminoeidae). Invertebrate Systematics, 32: 1336–1387. 

Valdés, Á., Breslau, E., Padula, V., Schrödl, M., Camacho, Y., Malaquias, M. A. E., Alexander, J., Bottomley, M., Vital, X., and Gosliner, T. M. 2018. Molecular and morphological systematics of DolabriferaGray, 1847 (Mollusca, Gastropoda, Heterobranchia, Aplysiomorpha). Zoological Journal of the Linnean Society, 184: 31–65. 

Moles, J., Ávila, C. and Malaquias, M. A. E. 2018. Systematic revision of the Antarctic gastropod family Newnesiidae (Heterobranchia: Cephalaspidea) with the description of a new genus and a new abyssal species. Zoological Journal of the Linnean Society, 183: 763–775. 

Bharate, M., Oskars, T, Malaquias, M. A. E. 2018. Diniatys callosa (Preston, 1908), new combination name for Haminoea callosafrom the Andaman Islands (India). Journal of Conchology, 43(1): 1–3.

Lobo-da-Cunha, A., Alves, Â., Oliveira, E. and Malaquias, M. 2018. Histological, histochemical and ultrastructural investigation of the male copulatory apparatus of Haminoea navicula(Gastropoda, Cephalaspidea). Journal of Morphology, 279(4): 554–565.  

Zamora-Silva, A. & Malaquias, M. A. E. 2018. Molecular phylogeny of the Aglajidae head-shield sea slugs (Heterobranchia: Cephalaspidea): new evolutionary lineages revealed and proposal of a new classification. Zoological Journal of the Linnean Society, 183(1): 1–51. 

Oskars,T. R., Mifsud, C., & Malaquias, M. A. E. 2017. Redescription of the Cephalaspidea gastropod Atys jeffreysi(Weinkauff, 1866) (Haminoeidae), with a discussion on the phylogenetic affinities of the Mediterranean species of the genus. Journal of Natural History, 51: 1593–1608. 

Tibiriça, Y. & Malaquias, M. A. E. 2017. The bubble snails (Gastropoda, Heterobranchia) of Mozambique: an overlooked biodiversity hotspot. Marine Biodiversity, 47(3), 791-811.

Malaquias, M. A. E., Zamora-Silva, A., Vitale, D., Spinelli. A., De Matteo, S., Giacobbe, S., Ortigosa, D. & Cervera J. L. 2017. The Suez Canal as a revolving door for marine species: a reply to Galil et al. (2016). Aquatic Invasions,&�Բ�����;12(1):&�Բ�����;1–4.&�Բ�����;

Garabedian, K., Malaquias, M.A.E., Crocetta, F., Zenetos, A., Kavadas, S. & Valdés, Á. 2017.Haminoea orteai Talavera, Murillo and Templado, 1987 (Mollusca: Gastropoda: Heterobranchia: Cephalaspidea), a widespread species in the Mediterranean and northeastern Atlantic. Cahiers de Biologie Marine, 58(1): 107-113. 

Malaquias, M. A. E., Ohnheiser, L. T., Oskars, T. R. & Willassen, E. 2017. Diversity and systematics of philinid snails (Gastropoda: Cephalaspidea) in West Africa with remarks on the biogeography of the region. Zoological Journal of the Linnean Society, 180(1): 1-35. 

Cruz-Rivera, E. & Malaquias, M. A. E. 2016. Ecosystem alterations and species range shifts: An Atlantic-Mediterranean cephalaspidean gastropod in an inland Egyptian lake. PLoS ONE, 11(6): e0156760.

Malaquias, M. A. E, Madrenas, E, & Ballesteros, M. 2016. First occurrence of the tropical Indo-West Pacific head-shield sea slug Chelidonura fulvipunctata in the Balearic Islands confirms its range extension into the western Mediterranean Sea. Mediterranean Marine Science 17 (New Mediterranean Biodiversity Records, March 2016): 232. 

Malaquias, M. A. E., Zamora-Silva, A., Vitale, D., Spinelli. A., De Matteo, S., Giacobbe, S., Ortigosa, D. & Cervera J. L. 2016. The Mediterranean Sea as a gateway for invasion of the Red Sea: The case of the Indo-West Pacific head-shield slug Chelidonura fulvipunctata Baba, 1938. Aquatic Invasions, 11(3): 247–255.

Padula, V., Bahia, J., Stöger, I., Camacho-García, Y., Malaquias, M. A. E., Cervera, J. L., & Schrödl, M. 2016. A test of color-based taxonomy in nudibranchs: molecular phylogeny and species delimitation of the Felimida clenchi (Mollusca: Chromodorididae) species complex.Molecular Phylogenetics and Evolution, 103: 215–229.

Kocot KM, Struck TH, Merkel J, Waits DS, Todt C, Brannock PM, Weese DA, Cannon JT, Moroz LL, Lieb B, Halanych KM. 2016. Phylogenomics of Lophotrochozoa with consideration of systematic error. Systematic Biology, in press.

Passos FD, Corrêa PVF, Todt C. 2016. A new species of Falcidens (Mollusca, Aplacophora, Caudofoveata) from the southeastern Brazilian coast: external anatomy, distribution, and comparison with Falcidens caudatus (Heath, 1918) from the USA. Marine Biodiversity, in press. 

Bergmeier FS, Haszprunar G, Todt C, Jörger KM. 2016. Lost in a taxonomic Bermuda Triangle: comparative 3D-microanatomy of cryptic mesopsammic Solenogastres (Mollusca). Organisms, Diversity & Evolution, in press.

Willassen, E., Williams, A. B., & Oskars, T. R. (2016). New observations of the enigmatic West African Cellana limpet (Mollusca: Gastropoda: Nacellidae).Marine Biodiversity Records, 9(1), 60.

Zamora, B. A. & Malaquias, M. A. E. 2016. Diet preferences of the Aglajidae: a family of cephalaspidean gastropod predators on tropical and temperate shores. Journal of the Marine Biological Association of the United Kingdom, 96(5): 1101–1112.

Caballer, M., Ortea J., Rivero, N., Carias, G., Malaquias, M. A. E. & Narciso, S. 2015. The opisthobranch gastropods (Mollusca: Heterobranchia) from Venezuela: an annotated and illustrated inventory of species. Zootaxa, 4034: 201–256.

Eilertsen, M. E. & Malaquias, M. A. E. 2015. Speciation in the dark: diversification and biogeography of the deep-sea gastropod genus Scaphander in the Atlantic Ocean. Journal of Biogeography, 42: 843–855.

Scherholz M, Redl E, Wollesen T, Todt C, Wanninger A. 2015. From complex to simple: myogenesis in an aplacophoran mollusk reveals key traits in aculiferan evolution. BMC Evolutionary Biology, 15: 201.

Wollesen T, Rodríguez Monje SV, Todt C, Degnan BM, Wanninger A. 2015. Ancestral role of Pax2/5/8 in molluscan
brain and multimodal sensory system development. BMC Evolutionary Biology, 15: 231.

Oskars, T. R., Bouchet, P., & Malaquias, M. A. E. 2015. A new phylogeny of the Cephalaspidea (Gastropoda: Heterobranchia) based on expanded taxon sampling and gene markers. Molecular Phylogenetics and Evolution, 89: 130–150.